Skip to main content
University of California Press

About the Book

Shorebirds are model organisms for illustrating the principles of ecology and excellent subjects for research. Their mating systems are as diverse as any avian group, their migrations push the limits of endurance, and their foraging is easily studied in the open habitats of estuaries and freshwater wetlands. This comprehensive text explores the ecology, conservation, and management of these fascinating birds. Beginning chapters examine phylogenetic relationships between shorebirds and other birds, and cover shorebird morphology, anatomy, and physiology. A section on breeding biology looks in detail at their reproductive biology. Because shorebirds spend much of their time away from breeding areas, a substantial section on non-breeding biology covers migration, foraging ecology, and social behavior. The text also covers shorebird demography, population size, and management issues related to habitat, predators, and human disturbances. Throughout, it emphasizes applying scientific knowledge to the conservation of shorebird populations, many of which are unfortunately in decline.

About the Author

Mark A. Colwell, Professor in the Wildlife Department at Humboldt State University, has been studying shorebirds for nearly thirty years.

Table of Contents

Preface and Acknowledgments

Part I Evolutionary Relationships, Anatomy and Morphology, and Breeding Biology

1 INTRODUCTION

Diversity and Distribution
Varied Ecomorphology
Diverse Social Systems
Globe- Trotting Migrants
Wetland Dependence
Conservation and Management
Rationale for and Organization of this book

2 SYSTEMATICS, PHYLOGENY, AND PHYLOGEOGRAPHY

Fossil History
A Brief History of Shorebird Systematics
Phylogeography
Hybridization in Shorebirds
Biogeography and Communities
Conservation Implications

3 MORPHOLOGY, ANATOMY, AND PHYSIOLOGY

Skeletal and Muscle System
Integumentary System
Sensory Apparatus, Foraging, and Digestion
Digestive System
Energetics and Thermoregulation
Osmoregulation
Reproductive System
Conservation Implications

4 MATING SYSTEMS

Defining a Mating System
The Role of Ecological Factors
Social versus Gene tic
Relationships
Parental Care Patterns
Evolution of Polyandry
Variance in Reproductive Success
Size Dimorphism
Sex Ratios
Conservation Implications


5 BREEDING BIOLOGY

Philopatry, Breeding Site Fidelity, and Dispersal
Spring Arrival Schedules
Courtship Behavior
Breeding Densities
Selection of a Breeding Site
Eggs
Incubation
Hatching
Chick Growth and Development
Conservation Implications

Part II Nonbreeding Ecology and Demography

6 MIGRATION

Origins and Evolution
Migration Strategies
Physiology of Migration
Hop, Skip, and Jump
Populations and Flyways
Conservation Implications

7 FORAGING ECOL OGY AND HABITAT USE

Diets
Foraging Maneuvers and Habitat Use
Acquiring Energy
Food Availability
Individual Variation
Conservation Implications

8 SHOREBIRDS AS PREDATORS

Shorebird Predators and Their Prey
Predicting Wetland Use
Competition and Food Limitation
Prey Reduction
Community Ecology
Conservation Implications

9 SPATIAL ECOLOGY AND WINTER SOCIAL ORGANIZATION

Quantifying Spatial Distributions
A Range of Social Organization
Roosts
Conservation Implications

10 POPULATION BIOLOGY

Demography
Survival
Productivity
Population Sizes and Trends
Monitoring Programs
Limiting Factors
Human Impacts
Conservation Implications

Part III Management and Conservation

11 HABITAT CONSERVATION AND MANAGEMENT

Decision Making in Wildlife Management
Wetland Conservation
Conservation Planning and Implementation
Wetland Management
Agricultural Lands
Salt Ponds
Sandy, Ocean- Fronting Beaches
Conservation Implications

12 MANAGING PREDATORS

Ethical Considerations and Decision Making
Do Predators Limit Shorebird Populations?
Methods of Control
Conservation Implications

13 MANAGING HUMAN DISTURBANCE

Definitions of Human Disturbance
Characterizing Disturbance
Responses to Disturbance
Managing Disturbance
Conservation Implications

14 EDUCATION AND OUTREACH

Professional Groups
Environmental Education
Ecotourism and Birding Festivals
Books and Online Resources
Conservation Implications

Appendix
Index



Reviews

“This comprehensive text explores the ecology, conservation, and management of these fascinating birds.”
The Guardian
“A fine book.”
Choice
“This is a splendid book, and there are lots of reasons why. First, the text is based on careful research into various aspects of shorebird biology worldwide. This excellent work should be on the shelf of anyone interested in shorebirds. . . . it is an extremely valuable source of information.”
Ibis
“[This book] will fill an important need for information on a group of birds whose existence is being threatened. . . . Recommended.”
Choice
“A pleasure to read and a real asset to a biologist’s bookshelf; both as a reference and as a source of inspiration.”
Emu-Austral Ornithology
“It is not often that one picks up a book and realizes he . . . has been missing it for years without realizing it. . . . Anyone considering getting into shorebirds, or taking on a student interested in doing so, will want a copy, or several copies of the book.”
The Condor
“Excellent . . . .Comprehensive.”
Northeastern Naturalist
"Anyone seeking a current and comprehensive reference on the ecology and conservation of shorebirds will find Colwell's effort outstanding. It provides a sorely needed companion to the numerous identification guides that have been published recently for this diverse and fascinating group of birds."—Robert Gill, Project Leader, Shorebird Research, USGS

"With their subtle beauty and amazing migrations, shorebirds continue to inspire and connect people across the planet. Amateur and professional shorebird enthusiasts will now be able to get the big picture from Colwell's authoritative and highly readable volume on a diverse and exciting group of birds, emblematic of many endangered habitats."—Theunis Piersma, University of Groningen