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University of California Press

About the Book

We all recognize the California Quail, with its striking good looks and mellow call notes. Widely distributed along the whole Pacific Coast, it has long been of interest to bird-lovers, sportsmen, ranchers, and wildlife biologists. In early times it was an important food source, and stocks were rapidly depleted by market hunters. Today its populations are being restored through the efforts of both private land-owners and government agencies. This comprehensive report summarizes what is known about the history, biology, and management of the species. It recounts the impact of settlement on quail, it describes the quail's intriguing social life, covey relationships, nesting behavior, and habitat requirements. Males, for example, play an important role in chick-rearing; chemical compounds in food substantially influence the nesting success of females. A major part of this book tells how to provide proper over, food, and drinking water that are needed to increase quail populations, whether for hunting or for observation and enjoyment. The suggestions offered stem from decades of scientific research and practical management effort. Written by a renowned wildlife biologist who worked on the quail for many years, this volume will be the definitive work from both the natural history and game management points of view. It is copiously illustrated with photographs and drawings, and will be of use to anyone who wishes to understand and foster the state bird.


We all recognize the California Quail, with its striking good looks and mellow call notes. Widely distributed along the whole Pacific Coast, it has long been of interest to bird-lovers, sportsmen, ranchers, and wildlife biologists. In early times it was a

About the Author

Aldo Starker Leopold (1913–1983) was an author, forester, zoologist and conservationist. He also served as professor at the University of California, Berkeley for thirty years. Throughout his life, Leopold was active in numerous wildlife and conservation groups throughout the United States.

Table of Contents

List of Figures and Tables
PREFACE
PROLOGUE
By Ian McMillan

PART I. THE BIRD AND ITS HISTORY

I. The Species and Its Distribution
California's State Bird
Native range of the species and its close relatives
Subspecies of California Quail
Mixing genetic strains through restocking
Extension of the California Quail range through transplants
The California Quail throughout the world
Competition with exotic game birds

2. Historical Perspectives
Indian use of California Quail
The quail in colonial California
Market hunting during the period of settlement
The quail "peak" 1860 to 1895
The quail decline

3. Early Efforts to Restore Quail in California
The genesis of management
Protective regulations
Quail refuges
Predator control
Artificial restocking
Habitat improvement
Resume of management experience

4. Land Use and Quail Habitat
Clear the valleys, graze the hills
Regional characteristics of quail range
Basic habitat needs
Cover
Food
Water
Changes in crops, tillage, and grazing practice in California
Quail in the chaparral
The invasion of brome grass in western rangelands
Logging in forested areas
Trends in land use

PART II. NATURAL HISTORY

5. Sociality in California Quail
Gregarious tendency in the species
Advantages in social organization
Sentry duty of cocks
Calls of the California Quail
Covey formation
Covey integrity
Covey territoriality
Covey size
Seasonal movements of quail

6. Covey Break-up and the Nesting Season
Weakening of the covey bond
Pairing behavior
The unmated cocks
Hormonal changes leading to reproduction
Development of the gonads
Changes in body weight
Nesting
Double-brooding
Non-breeding in unfavorable years
Quail reproduction in mesic and cool regions

7. Growth and Development of the Young Brooding
Nutrition of young chicks
The intestinal fauna
Rate of growth of young quail
Bursa of Fabricius
Plumages and molts
Plumage molt in adults
Sexual maturity in California Quail

8. Sex and Age Ratios and Their Interpretation
Sources of composition data
The annual cycle-!
Sex ratio
Age ratio
Year-to-year differences in production
Maximum age of quail

9. Rainfall as a Factor Affecting Reproductive Success
Rainfall and quail reproduction in northern and coastal ranges
Rainfall and quail reproduction in arid zones
Forb years vs. grass years
Pre-conditioning of breeding quail
Quail nutrition and reproduction
Green food as a stimulant to breeding in desert rodents
The possible role of steroids in regulating breeding
Legume seeds as a possible stimulus to breeding
Rainfall as a direct stimulant to breeding
Concomitant benefits of rainfall to quail reproduction
Relation of population density to breeding success

10. Quail Mortality
Mortality vs. natality
Quail predators
Disease and parasites
Agricultural poisons
Starvation
Hunting as a source of mortality
Miscellaneous losses
Density-dependent vs. density-independent mortality

PART III. QUAIL MANAGEMENT

11. Supplying Cover Needs
The strategy of managing quail range
The need for brush coverts
Brush clearing
Brush management
Planting escape cover
Roosting cover
Brush piles
Other types of cover
Nesting cover
The significance of cover in quail management

12. Making Food Available
Food as a limiting factor
Feeding habits of California Quail
Seasonal changes of diet
Green foods
Agricultural crop residues
Grazing in relation to food availability
Other cultural procedures to stimulate forb growth
Augmenting food in chaparral areas
Artificial feeding
The husbandry of quail range

13. Supplying Drinking Water
Is water necessary?
Moisture requirements of broods
Normal drinking habits of California Quail
Distance traveled to water
Need for cover near the water source
Artificial watering devices
The gallinaceous guzzler

14. Hunting the California Quail
Hunting philosophy
Hunting rights
Hunting strategy
The role of the dog
Reassembly of hunted coveys
Recording the results of quail hunting
Season and bag limits
The hunting kill of quail in Califomia
Predicting the quail crop from spring call counts
Allowable harvest
Does hunting endanger quail populations?

15. Backyard Quail
Quail for pleasure
Cover
Food
Water
Cats and dogs
A quail call at daylight

EPILOGUE

APPENDICES
A. Quail in Aboriginal California.
By Karen M. Nissen.
B. Foods of the California Quail.
By Bruce M. Browning.
C. Effects of Differing Rainfall on Breeding of California Quail
in an Arid Environment.
By Michael J. Erwin.

BIBLIOGRAPHY
INDEX

Reviews

"The avowed objective of the author of this book was "to assemble in one set of covers all that is known to date about the ecology, natural history, and management of the species." To the best of my knowledge, he has succeeded. . . . The book is highly readable, well organized, and will be of interest and value to laymen and professional wildlife managers alike."
Journal of Wildlife Management
"Very readable . . . an intensive, comprehensive review of the life history, population ecology, and management of this species that is clearly understandable to the ornithologist, game manager, and knowledgeable layman."
Journal of the West
"The combination of such an evocative species and an experienced wildlife biologist, like Starker Leopold, holds much promise, and the reader is not likely to be disappointed. This book is an outstanding example of a readable, yet scholarly and factual, life history account that will be of interest to not only professional ornithologists and wildlife managers, but also to amateur naturalists."
The Auk
"More than a single-species book; it is practical field ecology with multiple lessons for anyone interested in land and wildlife."
Ecology
"This will be the definitive work on this game species and its management."
Pacific Discovery
"Hopefully, this book will spark considerable interest among land managers throughout the range of the California Quail, and prompt them to set aside habitat for this species. The lessons provided in this book also have applicability throughout North America and beyond; one can substitute any species for the quail and apply the same principles expounded here equally well. The second section of the book will appeal to ornithologists in general in its summation of the salient data on the biology of this bird. The many, as yet unanswered, questions about "the natural controls over population numbers in this fascinating species" should stimulate us to further research efforts,"
Wilson Bulletin